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Mater is a tow truck voiced by Larry the Cable Guy. Along with the movie, he stars in a short, Mater and the Ghostlight, and acts as an announcer in the Cars video game. He speaks with a thick Southern accent with a bit of hillbilly twang, and often misspells words or uses incorrect syntax. Mater is also the self-proclaimed world champion backwards driver. He attributes his skill to his rear-view mirrors and his own philosophy. In the movie, Mater befriends and supports McQueen all throughout his adventure in Radiator Springs. He shows McQueen around the town, doing various things as "tractor tipping".

Mater is based on a 1951 International Harvester (IHC) Pickup truck, converted into a tow truck, but looks more like a mid-'50s Chevrolet. His color was originally baby blue, but over the years, he has rusted into a light brown/orange color, with some of the blue still showing through. He is missing his hood and has only a single headlight (his right). His license plate number is A113 (this is a recurring hidden element in Pixar's movie). His final personality was based on his voice actor, Larry the Cable Guy, and he uses many of Larry's catchphrases, including "Git-R-Done" during the final race sequence and "I don't care who you are; that's funny right there" during the tractor tipping scene, among others.

Mater also appears with his buddy Lightning at Disney's California Adventure of Disneyland Resort in Anaheim, CA and Disney's MGM Studios of Walt Disney World in Lake Buena Vista, FL for meet and greets. The 2 friends are very unique characters because they are real vehicles instead of others.